Critical Total Progeny Has a Heavy Tail
In a critical branching process, each individual has one child on average. That balance does not mean that population sizes remain near one: the process eventually dies out, but occasionally it produces a large family tree. When the offspring variance is finite and positive, the probability that the entire tree contains exactly \(n\) individuals decays at a power-law rate proportional to \(n^{-3/2}\). The lattice span of the offspring distribution determines the leading constant and which tree sizes are possible.
Work on a probability space \((\Omega,\mathcal{F},\mathbb{P})\). Let \((Z_n)_{n\ge0}\) be a Galton–Watson branching process, in discrete generation time \(n\), with countable state space \(S=\{0,1,2,\ldots\}\), initial population \(Z_0=1\), and offspring variable \(\xi\). Each individual independently produces a nonnegative integer number of children distributed as \(\xi\). Assume throughout that \(\mathbb{E}\xi=1\) and \(0<\operatorname{Var}(\xi)=\sigma^2<\infty\). These assumptions exclude the deterministic one-child process.
Write \(\xi_1,\ldots,\xi_n\) for independent copies of \(\xi\), and \(S_n=\xi_1+\cdots+\xi_n\). The earlier material on Dwass’s formula gives, for every integer \(n\ge1\),
This identity reduces a branching-tree probability to a sum of independent offspring counts. Here the sum has mean \(n\), while the required value \(n-1\) is one below its mean. The local central limit theorem evaluates the probability of this single lattice value.
The Lattice Local Limit Step
For the branching process, take \(\mu=1\) and \(x=n-1\). This value is in the required lattice precisely when \(n-1\equiv na\pmod d\). Call such an \(n\) lattice-admissible. At an admissible \(n\), the centered, scaled target is \(-1/(\sigma\sqrt{n})\), which tends to zero. The local central limit theorem therefore gives
The theorem also implies that this probability is positive for all sufficiently large lattice-admissible \(n\). Congruence is necessary for attainability at every \(n\), but it need not be sufficient at small \(n\).
Proof. Dwass’s formula applies to the total progeny of a Galton–Watson process started from one individual, so \(\mathbb{P}(T=n)=n^{-1}\mathbb{P}(S_n=n-1)\). At an admissible \(n\), the lattice local central limit theorem applies because the offspring variables are identically distributed, have mean \(1\), finite positive variance \(\sigma^2\), and span \(d\). The target \(n-1\) differs from the mean \(n\) by \(-1\), so its standardized displacement tends to zero. Hence \(\mathbb{P}(S_n=n-1)=d(\sigma\sqrt{2\pi n})^{-1}(1+o(1))\). Dividing by \(n\) proves the stated asymptotic. If \(n\) is inadmissible, \(S_n\) lies in \(na+d\mathbb{Z}\) and \(n-1\) does not, so the sum probability and, by Dwass’s formula, the total-progeny probability are both zero. \(\square\)
Worked Examples
Worked Example: Binary Offspring and Odd Total Sizes
Suppose \(\mathbb{P}(\xi=0)=\mathbb{P}(\xi=2)=\tfrac12\). The mean is \(0\cdot\tfrac12+2\cdot\tfrac12=1\). Also, \(\mathbb{E}[\xi^2]=0^2\cdot\tfrac12+2^2\cdot\tfrac12=2\), so \(\sigma^2=2-1^2=1\). The support differences are \(2\) and \(-2\), giving span \(d=2\).
For a support point \(a=0\), the sum \(S_n\) is always even. The target \(n-1\) is even exactly when \(n\) is odd. Thus even total sizes have probability zero. At \(n=3\), the target sum is \(2\). Exactly one of the three offspring values must equal \(2\), so
Check: A tree of total size \(3\) must have a root with two children, each of whom has no children. Its probability is \(\tfrac12\cdot\tfrac12\cdot\tfrac12=\tfrac18\), agreeing with Dwass’s formula. Along odd \(n\), the asymptotic constant is \(d/(\sigma\sqrt{2\pi})=2/\sqrt{2\pi}\). The factor \(2\) matters: the asymptotic is along the odd integers, while the probabilities at even integers are exactly zero.
Worked Example: Poisson Offspring
Let \(\xi\) have a Poisson distribution with parameter \(1\): \(\mathbb{P}(\xi=k)=e^{-1}/k!\) for \(k=0,1,\ldots\). Its mean and variance are both \(1\), and its support has span \(d=1\). The sum \(S_n\) is Poisson with parameter \(n\). Dwass’s formula gives, for every \(n\ge1\),
For example, at \(n=2\), this is \(e^{-2}\cdot 2^1/2!=e^{-2}\). Directly, a tree of size \(2\) requires the root to have one child and that child to have none, with probability \((e^{-1})(e^{-1})=e^{-2}\).
Stirling’s formula, \(n!\sim\sqrt{2\pi n}(n/e)^n\), yields
Check: The general constant is \(d/(\sigma\sqrt{2\pi})=1/\sqrt{2\pi}\), exactly the constant obtained from the explicit formula.
Worked Example: Congruence Need Not Ensure Small-Size Attainability
Consider the offspring law \(\mathbb{P}(\xi=0)=0.56=\tfrac{14}{25}\), \(\mathbb{P}(\xi=2)=0.40=\tfrac{10}{25}\), and \(\mathbb{P}(\xi=5)=0.04=\tfrac{1}{25}\). These probabilities sum to \(\tfrac{14+10+1}{25}=1\), and the mean is \(2\cdot\tfrac{10}{25}+5\cdot\tfrac{1}{25}=1\). The second moment is \(4\cdot\tfrac{10}{25}+25\cdot\tfrac{1}{25}=\tfrac{65}{25}=\tfrac{13}{5}\), so the variance is \(\tfrac{13}{5}-1=\tfrac85>0\). The support differences have greatest common divisor \(1\), so \(d=1\).
Every integer \(n\) satisfies the lattice congruence when \(d=1\). But for \(n=2\), the Dwass target is \(S_2=1\), and a sum of two values from \(\{0,2,5\}\) cannot equal \(1\). Therefore \(\mathbb{P}(T=2)=0\). Equivalently, a tree of size \(2\) would require the root to have exactly one child, which this offspring law does not allow.
Check: Dwass’s formula gives \(\mathbb{P}(T=2)=\tfrac12\mathbb{P}(S_2=1)=0\). The local central limit theorem’s positive approximation concerns sufficiently large admissible \(n\); it does not assert positivity at every small \(n\).
Extinction Is Certain, but the Mean Total Progeny Is Infinite
The tail asymptotic describes the size distribution, but almost-sure extinction is a separate fact. Let \(G(s)=\mathbb{E}[s^\xi]\), \(0\le s\le1\), be the offspring probability generating function. Because the offspring mean is \(1\) and the variance is positive, the offspring law is not concentrated at \(1\). It must have positive probability of zero offspring and positive probability of at least two offspring: if \(\xi\ge1\) almost surely, mean \(1\) would force \(\xi=1\) almost surely; and if \(\xi\le1\) almost surely, mean \(1\) would do the same.
Proof. Let \(q_n=\mathbb{P}(Z_n=0)\). Since zero is absorbing, this is also the probability of extinction by generation \(n\). Independence of offspring gives \(q_0=0\) and \(q_{n+1}=G(q_n)\). The sequence \(q_n\) is nondecreasing and bounded above by \(1\), so it has a limit \(q\). Continuity of \(G\) on \([0,1]\) gives \(q=G(q)\). Also \(G(1)=1\) and \(G'(1)=\mathbb{E}\xi=1\). Since \(\mathbb{P}(\xi\ge2)>0\), \(G\) is strictly convex on \((0,1)\); its tangent at \(1\) is the line \(s\mapsto s\). Thus \(G(s)>s\) for \(0\le s<1\), so \(1\) is its only fixed point in \([0,1]\). Consequently \(q=1\), proving extinction almost surely. For the expectation, let \(\mathcal{F}_n\) contain the population and reproduction information through generation \(n\). Conditional on \(\mathcal{F}_n\), the expected next generation is \(\mathbb{E}[Z_{n+1}\mid\mathcal{F}_n]=Z_n\mathbb{E}\xi=Z_n\). Induction from \(Z_0=1\) gives \(\mathbb{E}Z_n=1\) for every \(n\). By monotone convergence applied to the partial sums of \(T\), \(\mathbb{E}T=\sum_{n\ge0}\mathbb{E}Z_n=\sum_{n\ge0}1=\infty\). \(\square\)
There is no contradiction between these conclusions. Almost-sure extinction says that the realized tree has finitely many individuals with probability one; it does not bound the average size of that tree. Rare, very large trees make the expectation infinite. The tail gives the same warning: along the admissible sizes, \(n\mathbb{P}(T=n)\) is asymptotic to a positive constant times \(n^{-1/2}\), whose sum diverges. The generation-by-generation expectation argument proves infinite mean without requiring a separate summation over the lattice-admissible sizes.
Common pitfalls. First, omitting the lattice-span factor gives the wrong constant whenever \(d>1\). In the binary example, a span-one calculation would miss the factor \(2\), and it would fail to record that even sizes have probability zero. Second, lattice congruence is not a guarantee of attainability at every finite \(n\); it is the local limit theorem that gives positivity for sufficiently large admissible \(n\). Third, almost-sure extinction does not imply finite expected total progeny. In the critical finite-variance case, extinction is certain while \(\mathbb{E}T\) is infinite.
Check Your Understanding
Use the Dwass identity and check the lattice condition before applying the asymptotic.
- For offspring values \(0\) and \(2\), which total-progeny sizes are lattice-admissible, and what is the asymptotic constant along those sizes?
- For a critical offspring law with span \(3\) and variance \(4\), write the leading constant in terms of \(\pi\) along admissible sizes.
- In the Poisson-offspring example, compute \(\mathbb{P}(T=1)\) directly from Dwass’s formula.
- Why does the lattice local central limit theorem not imply that every congruence-admissible small \(n\) has positive total-progeny probability?
- Give the generation-by-generation calculation that proves \(\mathbb{E}T=\infty\), and explain why it does not contradict almost-sure extinction.